Photo of Paul Lewis

Paul Lewis


Paul is a Professor in the Ecology and Evolutionary Biology Department. He obtained his B.S. in Biology and Mathematics from Georgetown College in 1982, M.S. in Biology from Memphis State University in 1984, and Ph.D. in Plant Biology from Ohio State University in 1991. He was a postdoc at North Carolina State University with Bruce Weir, and at the Laboratory of Molecular Systematics, Smithsonian Institution, with David Swofford (now at Duke University). Paul came to UConn after 3 years as an Assistant Professor in the Biology Department at the University of New Mexico.


Suman Neupane

Suman Neupane

Ph.D. Student

Suman started working toward his Ph.D. here in January, 2013, after completing much of his dissertation research at Old Dominion University. He completed his Ph.D. in December, 2016, and continued on at UConn for another year as a postdoctoral researcher in the lab.

He studies flowering plants related to the genus Hedyotis (family Rubiaceae, the coffee family), which those in the eastern United States will recognize as the small blue flowers known as bluets carpeting roadsides and lawns in early spring. Hedyotis has a wide global distribution, however, and in addition to resolving the phylogeny and biogeography of this group Suman is interested in the tendency of Hedyotis to evolve secondary woodiness in tropical montane islands, which mimics the evolution of secondary woodiness common in oceanic island systems.

Suman is now a postdoctoral researcher in the Josef Uyeda laboratory at Virginia Tech. You can reach him at

Photo of Karolina Fučíková

Karolina Fučíková

Postdoctoral Researcher

Karolina Fučíková obtained her Ph.D. in the UConn EEB department (advisor L. Lewis) on the systematics of chlorophyte green algae, and later took the leading role in acquiring, characterizing and analyzing green algal chloroplast and mitochondrial genomes for the GrAToL (Assembling the Green Algal Tree of Life) and collaborated with other Lewis lab members and statisticians in the UConn Statistics department on the Bayesian Phylogenetic Information Content project.

Karolina is now an Assistant Professor in Biology at Assumption College in Worcester, Massachusetts. You can reach her at

Photo of Yu (Daniel) Fan

Yu Fan

Ph.D. Student

Daniel obtained his Ph.D. from UConn in 2011 with a dissertation entitled Bayesian phylogenetics, model selection, and methods of detecting non-independent and heterotachous molecular evolution. He obtained his undergraduate degree from the Ocean University of China in Qingdao, and Master’s degree in Molecular Systematics from the Institute of Oceanology, Chinese Academy of Sciences (IOCAS).

After leaving UConn, Daniel was a postdoctoral researcher in Wenyi Wang’s lab at M. D. Anderson Cancer Center in Houston, Texas.

He is currently at Philips Research North America in Cambridge, MA. You can reach him at

Mark Holder photo

Mark Holder

Associate Professor

Mark was a postdoctoral research associate in the lab for 2 years. Mark received his Ph.D. from the University of Texas, Austin, in 2001, and moved on to a postdoctoral position with David Swofford (then at Florida State University) in 2003. He is now an Associate Professor in the Department of Ecology and Evolutionary Biology at the University of Kansas. See Mark’s web site for more information.